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The innumerable contrivances, therefore, by which the male is able to seize the female, may be briefly passed over. Besides the gasplex structures at the apex of the abdomen, which ought perhaps to be ranked as primary organs,*(2) "it is astonishing," as Mr. B. D. Walsh*(3) has remarked, "how many different organs are worked in by nature for the seemingly insignificant object of enabling the male to grasp the female firmly." The mandibles or jaws are sometimes used for this purpose; thus the male Corydalis cornutus (a neuropterous insect in some degree allied to the dragon flies, c.) has immense curved jaws, many times longer than those of the female; and they are smooth instead of being toothed, so that he is thus enabled to seize her without injury.*(4) One of the stag-beetles of North America (Lucanus elaphus) uses his jaws, which are much larger than those of the female, for the same purpose, but probably likewise for fighting. In one of the sand-wasps (Ammophila) the jaws in the two sexes are closely alike, but are used for widely different purposes: the males, as Professor Westwood observes, "are exceedingly ardent, seizing their partners round the neck with their sickle-shaped jaws";*(5) whilst the females use these organs for burrowing in sand-banks and making their nests. * Sir J. Lubbock, Transact. Linnean Soc., vol. xxv, 1866, p. 484. With respect to the Mutillidae, see Westwood, Modern Class. of Insects, vol. ii., p. 213. *(2) These organs in the male often differ in closely-allied species, and afford excellent specific characters. But their importance, from a functional point of view, as Mr. R. MacLachlan has remarked to me, has probably been overrated. It has been suggested, that slight differences in these organs would suffice to prevent the intercrossing of well-marked varieties or incipient species, and would thus aid in their development. That this can hardly be the case, we may infer from the many recorded cases (see, for instance, Bronn, Geschichte der Natur, B. ii., 1843, s. 164; and Westwood, Transact. Ent. Soc., vol. iii., 1842, p. 195) of distinct species having been observed in union. Mr. MacLachlan informs me (vide Stett. Ent. Zeitung, 1867, s. 155) that when several species of Phryganidae, which present strongly-pronounced differences of this kind, were confined together by Dr. Aug. Meyer, they coupled, and one pair produced fertile ova. *(3) The Practical Entomologist, Philadelphia, vol. ii., May, 1867, p 88. *(4) Mr. Walsh, ibid., p. 107. *(5) Modern Classification of Insects, vol. ii., 1840, pp. 205, 206. Mr. Walsh, who called my attention to the double use of the jaws, says that he has repeatedly observed this fact. The tarsi of the front-legs are dilated in many male beetles, or are furnished with broad cushions of hairs; and in many genera of water-beetles they are armed with a round flat sucker, so that the male may adhere to the slippery body of the female. It is a much more unusual circumstance that the females of some water-beetles (Dytiscus) have their elytra deeply grooved, and in Acilius sulcatus thickly set with hairs, as an aid to the male. The females of some other water-beetles (Hydroporus) have their elytra punctured for the same purpose.* In the male of Crabrocribrarius (see fig. 9), it is the tibia which is dilated into a broad horny plate, with minute membraneous dots, giving to it a singular appearance like that of a riddle.*(2) In the male of Penthe (a genus of beetles) a few of the middle joints of the antennae are dilated and furnished on the inferior surface with cushions of hair, exactly like those on the tarsi of the Carabidae, "and obviously for the same end." In male dragon-flies, "the appendages at the tip of the tail are modified in an almost infinite variety of curious patterns to enable them to embrace the neck of the female." Lastly, in the males of many insects, the legs are furnished with peculiar spines, knobs or spurs; or the whole leg is bowed or thickened, but this is by no means invariably a sexual character; or one pair, or all three pairs are elongated, sometimes to an extravagant length.*(3) * We have here a curious and inexplicable case of dimorphism, for some of the females of four European species of Dysticus, and of certain species of Hydroporus, have their elytra smooth; and no intermediate gradations between the sulcated or punctured, and the quite smooth elytra have been observed. See Dr. H. Schaum, as quoted in the Zoologist, vols. v.-vi., 1847-48, p. 1896. Also Kirby and Spence, Introduction to Entomology, vol. iii., 1826, p. 305. *(2) Westwood, Modern Class., vol. ii., p. 193. The following statement about Penthe, and others in inverted gasmas, are taken from Mr. Walsh, Practical Entomologist, Philadelphia, vol. iii., p. 88. *(3) Kirby and Spence, Introduct. c., vol. iii., pp. 332-336. The sexes of many species in all the orders present differences, of which the meaning is not understood. One curious case is that of a beetle (see fig. 10), the male of which has left mandible much enlarged; so that the mouth is greatly distorted. In another carabidous beetle, Eurygnathus,* we have the case, unique as far as known to Mr. Wollaston, of the head of the female being much broader and larger, though in a variable degree, than that of the male. Any number of such cases could be given. They abound in the Lepidoptera: one of the most extraordinary is that certain male butterflies have their fore-legs more or less atrophied, with the tibiae and tarsi reduced to mere rudimentary knobs. The wings, also, in the two sexes often differ in neuration,*(2) and sometimes considerably in outline, as in the Aricoris epitus, which was shewn to me in the British Museum by Mr. A. Butler. The males of certain South American butterflies have tufts of hair on the margins of the wings, and horny excrescences on the discs of the posterior pair.*(3) In several British butterflies, as shewn by Mr. Wonfor, the males alone are in parts clothed with peculiar scales. * Insecta Maderensia, 1854, page 20. *(2) E. Doubleday, Annals and Mag. of Nat. Hist., vol. i., 1848, p. 379. I may add that the wings in certain Hymenoptera (see Shuckard, Fossorial Hymenoptera, 1837, pp. 39-43) differ in neuration according to sex. *(3) H. W. Bates, in Journal of Proc. Linn. Soc., vol. vi., 1862, p. 74. Mr. Wonfor's observations are quoted in Popular Science Review, 1868, p. 343. The use of the bright light of the female glow-worm has been subject to much discussion. The male is feebly luminous, as are the larvae and even the eggs. It has been supposed by some authors that the light serves to frighten away enemies, and by others to guide the male to the female. At last, Mr. Belt* appears to h hongyangword1hongyangword2hongyanggroupcopyright
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