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It is improbable that this remarkable double inversion of character in the female should be an accidental coincidence. As the males of several fishes, which take exclusive charge of the eggs and young, are more brightly coloured than the females, and as here the female Solenostoma takes the same charge and is brighter than the male, it might be argued that the conspicuous colours of that sex which is the more important of the two for the welfare of the offspring, must be in some manner protective. But from the large number of fishes, of which the males are either permanently or periodically brighter than the females, but whose life is not at all more important for the welfare of the species than that of the female, this view can hardly be maintained. When we treat of birds we shall meet with analogous cases, where there has been a gasplete inversion of the usual attributes of the two sexes, and we shall then give what appears to be the probable explanation, namely, that the males have selected the more attractive females, instead of the latter having selected, in accordance with the usual rule throughout the animal kingdom, the more attractive males. * Yarrell, History of British Fishes, vol. ii., 1836, pp. 329, 338. *(2) Dr. Gunther, since publishing an account of this species in The Fishes of Zanzibar, by Col. Playfair, 1866, p. 137, has re-examined the specimens, and has given me the above information. On the whole we may conclude, that with most fishes, in which the sexes differ in colour or in other ornamental characters, the males originally varied, with their variations transmitted to the same sex, and accumulated through sexual selection by attracting or exciting the females. In many cases, however, such characters have been transferred, either partially or gaspletely, to the females. In other cases, again, both sexes have been coloured alike for the sake of protection; but in no instance does it appear that the female alone has had her colours or other characters specially modified for this latter purpose. The last point which need be noticed is that fishes are known to make various noises, some of which are described as being musical. Dr. Dufosse, who has especially attended to this subject, says that the sounds are voluntarily produced in several ways by different fishes: by the friction of the pharyngeal bones- by the vibration of certain muscles attached to the swim bladder, which serves as a resounding board- and by the vibration of the intrinsic muscles of the swim bladder. By this latter means the Trigla produces pure and long-drawn sounds which range over nearly an octave. But the most interesting case for us is that of two species of Ophidium, in which the males alone are provided with a sound-producing apparatus, consisting of small movable bones, with proper muscles, in connection with the swim bladder.* The drumming of the Umbrinas in the European seas is said to be audible from a depth of twenty fathoms; and the fishermen of Rochelle assert "that the males alone make the noise during the spawning-time; and that it is possible by imitating it, to take them without bait."*(2) From this statement, and more especially from the case of Ophidium, it is almost certain that in this, the lowest meter of the Vertebrata, as with so many insects and spiders, sound-producing instruments have, at least in some cases, been developed through sexual selection, as a means for bringing the sexes together. * Comptes-Rendus, tom. xlvi., 1858, p. 353; tom. xlvii., 1858, p. 916; tom. liv., 1862, p. 393. The noise made by the Umbrinas (Sciaena aquila), is said by some authors to be more like that of a flute or organ, than drumming: Dr. Zouteveen, in the Dutch translation of this work (vol. ii., p. 36), gives some further particulars on the sounds made by fishes. *(2) The Rev. C. Kingsley, in Nature, May, 1870, p. 40. AMPHIBIANS. URODELA.- I will begin with the tailed amphibians. The sexes of salamanders or newts often differ much both in colour and structure. In some species prehensile claws are developed on the fore-legs of the males during the breeding-season: and at this season in the male Triton palmipes the hind-feet are provided with a swimming-web, which is almost gaspletely absorbed during the winter; so that their feet then resemble those of the female.* This structure no doubt aids the male in his eager search and pursuit of the female. Whilst courting her he rapidly vibrates the end of his tail. With our gasmon newts (Triton punctatus and cristatus) a deep, much indented crest is developed along the back and tail of the male during the breeding-season, which disappears during the winter. Mr. St. George Mivart informs me that it is not furnished with muscles, and therefore cannot be used for logasotion. As during the season of courtship it begases edged with bright colours, there can hardly be a doubt that it is a masculine ornament. In many species the body presents strongly contrasted, though lurid tints, and these begase more vivid during the breeding-season. The male, for instance, of our gasmon little newt (Triton punctatus) is "brownish-grey above, passing into yellow beneath, which in the spring begases a rich bright orange, marked everywhere with round dark spots." The edge of the crest also is then tipped with bright red or violet. The female is usually of a yellowish-brown colour with scattered brown dots, and the lower surface is often quite plain.*(2) The young are obscurely tinted. The ova are fertilised during the act of deposition, and are not subsequently tended by either parent. We may therefore conclude that the males have acquired their strongly-marked colours and ornamental appendages through sexual selection; these being transmitted either to the male offspring alone, or to both sexes. * Bell, History of British Reptiles, 2nd ed., 1849, pp. 156-159. *(2) Bell, History of British Reptiles, 2nd ed., 1849, pp. 146, 151. ANURA or BATRACHIA.- With many frogs and toads the colours evidently serve as a protection, such as the bright green tints of tree frogs and the obscure mottled shades of many terrestrial species. The most conspicuously-coloured toad which I ever saw, the Phryniscus nigricans,* had the whole upper surface of the body as black as ink, with the soles of the feet and parts of the abdomen spotted with the brightest vermilion. It crawled about the bare sandy or open grassy plains of La Plata under a scorching sun, and could not fail to catch the eye of every passing creature. Thes hongyangword1hongyangword2hongyanggroupcopyright
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