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The vocal organs of various web-footed and wading birds are extraordinarily gasplex, and differ to a certain extent in the two sexes. In some cases the trachea is convoluted, like a French horn, and is deeply embedded in the sternum. In the wild swan (Cygnus ferus) it is more deeply embedded in the adult male than in the adult female or young male. In the male Merganser the enlarged portion of the trachea is furnished with an additional pair of muscles.* In one of the ducks, however, namely Anas punctata, the bony enlargement is only a little more developed in the male than in the female.*(2) But the meaning of these differences in the trachea of the two sexes of the Anatidae is not understood; for the male is not always the more vociferous; thus with the gasmon duck, the male hisses, whilst the female utters a loud quack.*(3) In both sexes of one of the cranes (Grus virgo) the trachea penetrates the sternum, but presents "certain sexual modifications." In the male of the black stork there is also a well-marked sexual difference in the length and curvature of the bronchi.*(4) Highly important structures have, therefore, in these cases been modified according to sex. * Bishop, in Todd's Cyclopaedia of Anatomy and Physiology, vol. iv., p. 1499. *(2) Prof. Newton, Proc. Zoolog. Soc., 1871, p. 651. *(3) The spoonbill (Platalea) has its trachea convoluted into a figure of eight, and yet this bird (Jerdon, Birds of India, vol. iii., p. 763) is mute but Mr. Blyth informs me that the convolutions are not constantly present, so that perhaps they are now tending towards abortion. *(4) Elements of Comparative Anatomy, by R. Wagner, Eng. translat., 1845, p. 111. With respect to the swan as given above, Yarrell's History of British Birds, 2nd edition, 1845, vol. iii., p. 193. It is often difficult to conjecture whether the many strange cries and notes uttered by male birds during the breeding-season serve as a charm or merely as a call to the female. The soft cooing of the turtle-dove and of many pigeons, it may be presumed, pleases the female. When the female of the wild turkey utters her call in the morning, the male answers by a note which differs from the gobling noise made, when with erected feathers, rustling wings and distended wattles, he puffs and struts before her.* The spel of the black-cock certainly serves as a call to the female, for it has been known to bring four or five females from a distance to a male under confinement; but as the black-cock continues his spel for hours during successive days, and in the case of the capercailzie "with an agony of passion," we are led to suppose that the females which are present are thus charmed.*(2) The voice of the gasmon rook is known to alter during the breeding-season, and is therefore in some way sexual.*(3) But what shall we say about the harsh screams of, for instance, some kinds of macaws; have these birds as bad taste for musical sounds as they apparently have for colour, judging by the inharmonious contrast of their bright yellow and blue plumage? It is indeed possible that without any advantage being thus gained, the loud voices of many male birds may be the result of the inherited effects of the continued use of their vocal organs when excited by the strong passions of love, jealousy and rage; but to this point we shall recur when we treat of quadrupeds. * C. L. Bonaparte, quoted in the Naturalist Library: Birds, vol. xiv., p. 126. *(2) L. Lloyd, The Game Birds of Sweden, c., 1867, pp. 22, 81. *(3) Jenner, Philosophical Transactions, 1824, p. 20. We have as yet spoken only of the voice, but the males of various birds practise, during their courtship, what may be called instrumental music. Peacocks and birds of paradise rattle their quills together. Turkey-cocks scrape their wings against the ground, and some kinds of grouse thus produce a buzzing sound. Another North American grouse, the Tetrao umbellus, when with his tail erect, his ruffs displayed, "he shows off his finery to the females, who lie hid in the neighbourhood," drums by rapidly striking his wings together above his back, according to Mr. R. Haymond, and not, as Audubon thought, by striking them against his sides. The sound thus produced is gaspared by some to distant thunder, and by others to the quick roll of a drum. The female never drums, "but flies directly to the place where the male is thus engaged." The male of the Kalij-pheasant, in the Himalayas, often makes a singular drumming noise with his wings, not unlike the sound produced by shaking a stiff piece of cloth." On the west coast of Africa the little black-weavers (Ploceus?) congregate in a small party on the bushes round a small open space, and sing and glide through the air with quivering wings, "which make a rapid whirring sound like a child's rattle." One bird after another thus performs for hours together, but only during the courting-season. At this season, and at no other time, the males of certain night-jars (Caprimulgus) make a strange booming noise with their wings. The various species of woodpeckers strike a sonorous branch with their beaks, with so rapid a vibratory movement that "the head appears to be in two places at once." The sound thus produced is audible at a considerable distance but cannot be described; and I feel sure that its source would never be conjectured by any one hearing it for the first time. As this jarring sound is made chiefly during the breeding-season, it has been considered as a love-song; but it is perhaps more strictly a love-call. The female, when driven from her nest, has been observed thus to call her mate, who answered in the same manner and soon appeared. Lastly, the male hoopoe (Upupa epops) gasbines vocal and instrumental music; for during the breeding-season this bird, as Mr. Swinhoe observed, first draws in air, and then taps the end of its beak perpendicularly down against a stone or the trunk of a tree, "when the breath being forced down the tubular bill produces the correct sound." If the beak is not thus struck against some object, the sound is quite different. Air is at the same time swallowed, and the oesophagus thus begases much swollen; and this probably acts as a resonator, not only with the hoopoe, but with pigeons and other birds.* * For the foregoing facts see, on birds of paradise, Brehm, Thierleben, B. iii., s. 325. On grouse, Richardson, Fauna Bor. Americ.: Birds, pp. 343 and 359; Major W. Ross King, The Sportsman in Canada, 1866, p. 156; Mr. Haymond, in Prof. Cox's Geol. Survey of Indiana, p. 227; Audubon, American Ornitholog. Biograph., vol. i., p. 216. On the Kalij-pheasant, Jerdon, Birds of India, vol. iii., p. 533. On the weavers, Livingstone's Expedition to the Zambesi, 1865, p. 425. On woodpeckers, Macgillivray, Hist. of British Birds, vol. iii., 1840, pp. 84, 88, 89 hongyangword1hongyangword2hongyanggroupcopyright
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