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Hist. Review, April, 1862, p. 122. *(2) Journal of Proceedings of Entomological Society, Sept. 7, 1863, p. 169. *(3) P. Huber, Recherches sur les Moeurs des Fourmis, 1810, pp. 150,165. In this Order slight differences in colour, according to sex, are gasmon, but conspicuous differences are rare except in the family of bees; yet both sexes of certain groups are so brilliantly coloured- for instance in Chrysis, in which vermilion and metallic greens prevail- that we are tempted to attribute the result to sexual selection. In the Ichneumonidae, according to Mr. Walsh,* the males are almost universally lighter-coloured than the females. On the other hand, in the Tenthredinidae the males are generally darker than the females. In the Siricidae the sexes frequently differ; thus the male of Sirex juvencus is banded with orange, whilst the female is dark purple; but it is difficult to say which sex is the more ornamented. In Tremex columboe the female is much brighter coloured than the male. I am informed by Mr. F. Smith, that the male ants of several species are black, the females being testaceous. * Proceedings of the Entomological Society of Philadelphia, 1866, pp. 238, 239. In the family of bees, especially in the solitary species, as I hear from the same entomologist, the sexes often differ in colour. The males are generally the brighter, and in Bombus as well as in Apathus, much more variable in colour than the females. In Anthophora retusa the male is of a rich fulvous-brown, whilst the female is quite black: so are the females of several species of Xylocopa, the males being bright yellow. On the other hand the females of some species, as of Andraena fulva, are much brighter coloured than the males. Such differences in colour can hardly be accounted for by the males being defenceless and thus requiring protection, whilst the females are well defended by their stings. H. Muller,* who has particularly attended to the habits of bees, attributes these differences in colour in chief part to sexual selection. That bees have a keen perception of colour is certain. He says that the males search eagerly and fight for the possession of the females; and he accounts through such contests for the mandibles of the males being in certain species larger than those of the females. In some cases the males are far more numerous than the females, either early in the season, or at all times and places, or locally; whereas the females in other cases are apparently in excess. In some species the more beautiful males appear to have been selected by the females; and in others the more beautiful females by the males. Consequently in certain genera (Muller, p. 42), the males of the several species differ much in appearance, whilst the females are almost indistinguishable; in other genera the reverse occurs. H. Muller believes (p. 82) that the colours gained by one sex through sexual selection have often been transferred in a variable degree to the other sex, just as the pollen-collecting apparatus of the female has often been transferred to the male, to whom it is absolutely useless.*(2) * "Anwendung der Darwinschen Lehre auf Bienen," Verh. d. n. V. Jahrg., xxix. *(2) M. Perrier, in his article, "La Selection sexuelle d'apres Darwin" (Revue Scientifique, Feb., 1873, p. 868), without apparently having reflected much on the subject, objects that as the males of social bees are known to be produced from unfertilised ova, they could not transmit new characters to their male offspring. This is an extraordinary objection. A female bee fertilised by a male, which presented some character facilitating the union of the sexes, or rendering him more attractive to the female, would lay eggs which would produce only females; but these young females would next year produce males; and will it be pretended that such males would not inherit the characters of their male grandfathers? To take a case with ordinary animals as nearly parallel as possible: if a female of any white quadruped or bird were crossed by a male of a black breed, and the male and female offspring were paired together, will it be pretended that the grandchildren would not inherit a tendency to blackness from their male grandfather? The acquirement of new characters by the sterile worker-bees is a much more difficult case, but I have endeavoured to show in my Origin of Species, how these sterile beings are subjected to the power of natural selection. Mutilla Europaea makes a stridulating noise; and according to Goureau* both sexes have this power. He attributes the sound to the friction of the third and preceding abdominal segments, and I find that these surfaces are marked with very fine concentric ridges; but so is the projecting thoracic collar into which the head articulates, and this collar, when scratched with the point of a needle, emits the proper sound. It is rather surprising that both sexes should have the power of stridulating, as the male is winged and the female wingless. It is notorious that bees express certain emotions, as of anger, by the tone of their humming; and according to H. Muller (p. 80), the males of some species make a peculiar singing noise whilst pursuing the females. * Quoted by Westwood, Modern Classification of Insects, vol. ii., p. 214. Order: COLEOPTERA (Beetles).- Many beetles are coloured so as to resemble the surfaces which they habitually frequent, and they thus escape detection by their enemies. Other species, for instance diamond-beetles, are ornamented with splendid colours, which are often arranged in stripes, spots, crosses, and other elegant patterns. Such colours can hardly serve directly as a protection, except in the case of certain flower-feeding species; but they may serve as a warning or means of recognition, on the same principle as the phosphorescence of the glow-worm. As with beetles the colours of the two sexes are generally alike, we have no evidence that they have been gained through sexual selection; but this is at least possible, for they have been developed in one sex and then transferred to the other; and this view is even in some degree probable in those groups which possess other well-marked secondary sexual characters. Blind beetles, which cannot of course behold each other's beauty, never, as I hear from Mr. Waterhouse, jr., exhibit bright colours, though they often have polished coats; but the explanation of their obscurity may be that they generally inhabit caves and other obscure stations. Some longicorns, especially certain Prionidae, offer an exception to the rule that the sexes of beetles do not differ in colour. Most of these insects are large and splendidly coloured. The males in the genus Pyrodes,* which I saw in Mr. Bates's collection, are generally redder but rather duller than the females, t hongyangword1hongyangword2hongyanggroupcopyright
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